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The son of a Japanese stewardess and an Italian pilot, Formichetti was born in Japan and grew up in Rome and Tokyo, attempting to assimilate in both cultures. As a child he trained as a classical pianist (and still plays for relaxation). But moving to London became his great desire and he used the excuse of attending an architecture school to which he was accepted in order to get there. Formichetti has explained that instead of attending he "literally walked in the front door of the architecture school and then ran out of the back one to go clubbing for three years." He became a student of the nightlife and street fashions, and began working at a fashionable boutique called 'The Pineal Eye' where he would become art director and head buyer for two years.
His collaboration with Lady Gaga as her stylist, or more appropriately fashion director, began in early May 2009 at a shoot with photographer Sebastian Faena for the July 2009 issue of V Magazine (V60), Formichetti saying that "It was an instant love." This was soon followed by a cover shoot for V61 (September issue) with Mario Testino. The following June he was fashion director for her performance at the 2009 MuchMusic Video Awards, and following that for the majority of Lady Gaga's important performances and appearances in the second half of 2009, including at the 2009 MTV Video Music Awards in September, where she appeared in four dresses by Jean-Paul Gaultier and one by Alexander McQueen, then on Saturday Night Live in October, where she appeared in a metal orb structure designed by Nazir Mazhar, and later at the Royal Variety Performance in December, where she appeared in a latex dress, by Atsuko Kudo, designed to resemble those worn by Elizabeth I of England, for Gaga's introduction to her namesake Elizabeth II. In the fall she also appeared in the third volume of Vogue Hommes Japan, the editorial titled "Lady Gaga & Araki's Tokyo Love" shot by Nobuyoshi Araki, with the fashion direction naturally by Formichetti. Their first collaboration in a music video together, the immensely successful Bad Romance, directed by Francis Lawrence, would later be voted Video of the Year and then win the Grammy Award for Best Short Form Music Video. In this Gaga appears in several outfits and footwear by Alexander McQueen, reappears in the orb by Nazir Mazhar, as well as in a "diamond crown" outfit by Franc Fernandez, and one by her creative team, the "Haus of Gaga" itself, among other pieces. Formichetti is also credited as stylist for her album The Fame Monster (shot by Hedi Slimane), and has been fashion director for her sold-out The Monster Ball Tour.
In late January 2010 he was stylist/fashion director for Lady Gaga's music video Telephone, directed by Jonas Åkerlund, which was later released in March. Immediately after this video was filmed came her appearance at the 52nd Grammy Awards (where Gaga performed with Elton John), for which Formichetti was also fashion director. In February he was again for her appearance at the 2010 BRIT Awards, where she dedicated her performance to their friend Alexander McQueen, who had committed suicide only five days earlier. Following their collaboration for several more appearances and performances over the next months came Formichetti's styling for her video Alejandro, directed by his collaborator in the past Steven Klein, and for the associated cover shoot by Terry Richardson for the mid-July issue of Rolling Stone. Her American Idol performance of "Alejandro" in early May was also styled by Formichetti. Among other projects, next came the styling for Richardson's shoot of Gaga (in a meat bikini) and Nick Knight's shoot of model Jo Calderone (Gaga in "drag") for the fifth volume of Vogue Hommes Japan. Around the same time Richardson shot Gaga as styled by Formichetti for the company Supreme NYC, but the shoot would wait until 2011 to be released both in the Supreme ad campaign and in the French magazine Purple. Knight then photographed her again, styled by Formichetti, for the upcoming September issue of Vanity Fair. In August the three would appear yet again in collaboration for the 30th birthday issue of the magazine i-D (the shoot itself actually dating from 2009), Formichetti also giving a short interview for the Gaga piece following hers. Then in September came her celebrated appearance, for which he did the styling, at the 2010 MTV Video Music Awards, where she wore (among other outfits) the famous "meat dress" by Franc Fernandez. Also in September the Fall issue of V Magazine (V67) appeared on newsstands, with Gaga, styled by Formichetti and shot by Mario Testino, on the cover with Marc Jacobs.

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After several months largely out of the spotlight (except on tour) Gaga returned in January 2011 to begin the serious promotion for her upcoming album Born This Way and Formichetti has been fashion director for the project. The music video for the title track "Born This Way" was directed later in January by Nick Knight, with Formichetti responsible for the styling. In February came her performance at the 53rd Grammy Awards where they collaborated with designer Hussein Chalayan on her appearance and performance. Gaga, interested in the concept of rebirth, sent Formichetti a picture of a human embryo, and he was first inspired to create, as he told her, "egg [shaped] shoes—like you were born that way, with eggs as feet." She took the idea further, resulting in the egg or "vessel" (designed by Chalayan) in which she was carried down the red carpet and later onto the stage, hatching out in a yolk-coloured latex outfit to perform her new single "Born This Way". She was wearing, as he tweeted, "Mugler by Nicola Formichetti." A "Born This Way"-associated shoot called "Alien Excess" done in January by photographer Mariano Vivanco for the Spring 2011 (March) issue of i-D features Gaga in a latex coat from the new Mugler menswear collection, with Formichetti responsible for the styling. A shot from the same session with Vivanco, but with different styling, also by Formichetti, appeared on the cover of Billboard in February.

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He has also done Gaga's styling (shot by Nick Knight) for her as spokesperson for the MAC VIVA GLAM Campaign of the MAC AIDS Fund. A new project called "The Masterpiece" has been announced in which Formichetti will collaborate with her for MAC to create a "wearable piece of art" composed of the profile photos of fans who join the campaign.
After months of rumours, in September 2010 Formichetti was announced as the new creative director for the Parisian fashion house founded by the celebrated designer Thierry Mugler (who has largely retired from the business), which he has renamed to MUGLER. He has said that it was in fact Lady Gaga who persuaded him to go for the job, when they were looking through pieces from the Mugler archive when shooting the video for "Telephone". At MUGLER Formichetti oversees designers Sébastien Peigné for womenswear and Romain Kremer for menswear. Joël Palix, director general of the company, said "We were looking for a young talent who could really bring energy to the brand... Nicola is a multiculural, techno-savvy expert involved in fashion, communication, image and entertainment. He and the appointed talented designers will represent a new direction for French fashion."

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The menswear, a collaboration with Romain Kremer, of Formichetti's debut (Fall/Winter) collection "Anatomy of Change" premiered during the Men's Paris fashion week on 19 January 2011. This was accompanied by a video shot by Mariano Vivanco, featuring Formichetti's new muse, the full-body tattooed Canadian model Rick Genest, and by a remix by Lady Gaga, as musical director for the menswear project, of a new track ("Scheiße") from her upcoming album Born This Way. The clothes, worn by Rico Genest and other models on the runway, recall those by Mugler himself from earlier decades, in such ways as the use of the colours navy blue and bright orange, or "hyper-modern"-style jackets lacking lapels. The collection also features some not-so-conventional latex pieces such as biker jackets, gloves, and even latex trousers. On the overall vision, Cathy Horyn of The New York Times describes Formichetti creating a "mechanical man" in the tradition of Mugler's fantastical woman, yet Adam Tschorn of the Los Angeles Times found plenty of the pieces surprisingly wearable. Formichetti himself says on the collection "I want to say something that has the past and the future, but what is really important to me is the present." Returning to Rico Genest, it was his discovery by Formichetti and then Gaga's urging which actually resulted in the menswear show, something not originally planned. Genest's discovery also influenced the collection itself. The show was streamed live from Paris on the Internet. Genest can be seen styled in MUGLER by Formichetti and shot by Mariano Vivanco in the sixth volume of Vogue Hommes Japan, the editorial being titled "Hard To Be Passive". In the Spring/Summer 2011 issue of GQ Style (UK) Formichetti is interviewed on the future of the brand, with Genest (also interviewed) shot in MUGLER by Karim Sadli for the editorial. On March 2 the womenswear, a collaboration with Sébastien Peigné, premiered. Lady Gaga was again musical director, previewing the track "Government Hooker" from her upcoming album as well as including the already released title track "Born This Way" in the soundtrack. But this time she actually performed on the runway in person for Formichetti's show, modelling pieces from his collection, called "Anatomy of Change Femme – Mode Sans Frontiers." Additionally, Rick Genest from the menswear collection made a reappearance on the runway among the models. The reaction of critics to the clothes was generally similar to their reaction to the menswear two months previously. Formichetti offers a silhouette in the Mugler tradition, "minimally rendered" as described by Cathy Horyn, and with sculptured shoulders (for the appropriate pieces), Alex Fury of SHOWstudio.com describing them as "architectonic". For the show Formichetti employed Rein Vollenga to contribute sculptural elements to the silhouette, Atsuko Kudo skin-like items of torn-up latex, and Franc Fernandez for Gaga's two hats. She herself gave input for the collection, associated with her Born This Way projects and performances, but is not financially involved with Formichetti's MUGLER. As he says, "It's just friends helping each other... I think when you start talking about money, it stops the whole creative process for me."